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Aiming at the traditional localization method usually aiming at the static wireless sensor network, can not suit the mobile sensor network with dynamic changing network structure and node distance, a locating and tracking method for mobile sensor node was proposed based on RSSI and improved MCL. Firstly, the RSSI ranging distance method and classic MCL was introduced, then an improved MCL algorithm was proposed by renewing the process of classic MCL such as predicating and filtering to five stages as anchor node broadcast its position using limit TTL flooding protocol, using Lagrange interpolation to predict the position and speed in the next time, obtaining anchor sampling area, particle filter and according the predicting position and speed in the next time and the current position to weight particle. The experiment was simulated in the MCL-Simulator shows the method in this paper can realize the node moving, compared with the other methods, it has less average locating error, so it has big feasibility.